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1.
In the present experiments, we examined the role of within-compound associations in the interaction of the overshadowing procedure
with conditioned stimulus (CS) duration, using a conditioned suppression procedure with rats. In Experiment 1, we found that,
with elemental reinforced training, conditioned suppression to the target stimulus decreased as CS duration increased (i.e.,
the CS duration effect), whereas, with compound reinforced training (i.e., the overshadowing procedure), conditioned suppression to the target stimulus
increased as CS duration increased. In subsequent experiments, we replicated these findings with sensory preconditioning and
demonstrated that extinction of the overshadowing stimulus results in retrospective revaluation with short CSs and in mediated
extinction with long CSs. These results highlight the role of the duration of the stimulus in behavioral control. Moreover,
these results illuminate one cause (the CS duration) of whether retrospective revaluation or mediated extinction will be observed. 相似文献
2.
Three experiments demonstrated that, following the extinction of an established conditioned stimulus (CS; e.g., tone), the
pairing of an orthogonal stimulus from another modality (e.g., light) with the unconditioned stimulus (US) results in strong
recovery of responding to the extinguished CS. This recovery occurred to about an equal degree regardless of whether or not
initial training contained unambiguous stimulus—reinforcer relationships—that is, consistent CS—US pairings—or some degree
of ambiguity, including intramodal discrimination training, partial reinforcement, or even cross-modal discrimination training
(tone vs. light). Experiments 1 and 2 demonstrated that this recovery of responding was largely specific to the extinguished
CS, but moderate generalization to other stimuli from the same modality did appear. The results are discussed with reference
to alternative mechanisms applicable to learning-dependent generalization between otherwise distinct CSs. These models assume
that such generalization is mediated by either a shared response, shared reinforcer, shared context, or shared hidden units
within a layered neural network. A specific layered network is proposed to explain the present results as well as other types
of savings seen previously in conditioning of the rabbit nictitating membrane response. 相似文献
3.
Matthew R. Tinsley William Timberlake Matthew Sitomer David R. Widman 《Learning & behavior》2002,30(3):217-227
Like other accounts of conditioned inhibition, behavior systems predicts (and Experiment 1 shows) that during summation and retardation tests, presentation of a negative conditioned stimulus (a CS−) created by discriminative Pavlovian food conditioning will interfere with a focal search response, such as nosing in the feeder. Unlike most other views, behavior systems predicts (and Experiment 2 shows) that the same CS− can potentiate a general search response, like attending to a moving artificial prey stimulus. Contacting the prey stimulus in extinction increased over baseline when a CS- but not a CS Novel preceded it. Experiment 3 showed this effect was not due to unconditioned qualities of the CS−. It appears that the effects of a discriminative CS-depend on the interaction of the training contingency with search modes related to the unconditioned stimulus (US), their perceptual-motor repertoires and environmental support, and the choice of response measure. 相似文献
4.
Philippe Oberling Adam S. Bristol Helena Matute Ralph R. Miller 《Learning & behavior》2000,28(2):172-186
Miller and Matute (1996) showed that blocking is attenuated when the blocked conditioned stimulus (CS) is “biologically significant” (i.e., when the CS has the potential to elicit vigorous responding of any kind). To the extent that blocking is representative of cue competition, this finding suggests that biological significance protects CSs against cue competition effects in general. In the present experiments, we tested this possibility by examining the influence of biological significance of CSs on other examples of cue competition, namely, overshadowing, the relative stimulus validity effect, and the degraded contingency effect in rats. In Experiment 1, we found that intense auditory stimuli induced transient unconditioned lick suppression, thereby indicating that intense sounds were of high inherent biological significance. In Experiment 2A, we found that cues with high inherent biological significance were protected from overshadowing. In Experiment 2B, this finding was extended to cues with high acquired biological significance, which was obtained through prior pairings with a reinforcer of the valence opposite to that used in the overshadowing treatment. In Experiments 3 and 4, we found that cues with high inherent biological significance attenuated the relative validity effect and the degraded contingency effect, respectively. These results lend support to the view that biological significance (inherent and acquired) protects stimuli from cue competition effects, a finding that is problematic for many contemporary theories of learning. 相似文献
5.
Four experiments used a within-subjects design with rats to study the effects of preexposure on the restoration of fear responses
(freezing) to an extinguished conditioned stimulus (CS). In each experiment, rats were preexposed to one CS (A), but not to
another (B), and then were exposed to pairings of each of these CSs with an aversive unconditioned stimulus (US). In each
experiment, there was less freezing to A than to B across extinction, showing a latent inhibitory effect of preexposure. There
was no differential recovery to A and B following either a US reexposure (Experiment 1) or a delay interval (Experiment 2). However, when a delay interval included US reexposure, there was greater recovery to the preexposed CS, A, than to the
nonpreexposed CS, B (Experiments 1, 3, and 4). These results suggest that the effects of US reexposure and delay combine to affect recovery from the depressive effects
of CS-alone exposure. The results are consistent with the view that US reexposure produces better mediated conditioning of
CSs that are strongly associated with the context. The results may additionally reflect an effect of preexposure on the learning
produced by extinction. 相似文献
6.
Adding limited female cues to a conditioned stimulus (CS) facilitates conditioned male sexual responding. In two experiments, we examined the mechanisms of this facilitation effect. The color of the female cues on the CS was varied in Experiment 1. Similarity between the CS plumage color and the color of the live female (the unconditioned stimulus [US]) could only partially account for the results. The extent to which the facilitation effect represents a specialization of sexual behavior was examined in Experiment 2 by comparing conditioning with either food or copulation as the US. The CSs with female cues elicited more approach and grab responses regardless of which US was used. However, uniquely sexual conditioned responses (mounts and cloacal contacts) were enhanced only when sexual reinforcement served as the US. These findings suggest that the facilitation effect of female cues represents a general feature of appetitive behavior systems. 相似文献
7.
In three experiments with rat subjects, we examined the effects of trial spacing in appetitive conditioning. Previous research
in this preparation suggests that self-generated priming of the conditional stimulus (CS) and/or unconditional stimulus (US)
in short-term memory is a cause of the trial-spacing effect that occurs with intertrial intervals (ITIs) of less than 240
sec. Experiment 1 nonetheless showed that a trial-spacing effect still occurs when ITIs are increased beyond 240 sec, and
that the effect of ITI over 60–1,920 sec on conditioned responding is best described as a linear function. In Experiment 2,
some subjects were removed from the context during the ITIs, preventing extinction of the context. Removal abolished the advantage
of the long ITI, suggesting the importance of exposure to the context during the long ITI. Experiment 3 still produced a trial-spacing
effect in a within-subjects design that controlled for the level of context conditioning and reinforcement rate in the absence
of the CS. Overall, the results are most consistent with the idea that adding time to the ITI above 240 sec facilitates conditioning
by extinguishing context-CS associations—and possibly context-US associations—that otherwise interfere with CS-US learning
through retrieval-generated priming (see, e.g., Wagner, 1981). 相似文献
8.
Rats were used in a lick suppression preparation to assess the contribution of conditioned-stimulus (CS)–context and context–unconditioned-stimulus
(US) associations to experimental extinction. Experiment 1 investigated whether strengthening the CS–acquisition context association enhances extinction by determining whether stronger
extinction is observed when CS-alone trials (i.e., extinction treatment) are administered in the acquisition context (AAC
renewal), relative to a context that is neutral with respect to the US (ABC renewal). Less recovery of responding to the CS
was observed in the former than in the latter case, extending the finding that AAC renewal is weaker than ABC renewal to our
lick suppression preparation. Experiment 2 assessed the contribution of the acquisition context–US association to extinction of a CS by examining the effect of postextinction
exposure to the acquisition context on responding to the extinguished CS. This manipulation enhanced responding to the extinguished
CS in AAC, but not ABC, renewal. Experiment 3 addressed the contribution of the CS–acquisition context association by examining the potential of a neutral stimulus, presented
in compound with the target CS during extinction treatment, to overshadow the CS–acquisition context association. This manipulation
enhanced responding to the extinguished CS in AAC, but not ABC, renewal. The results stress the important role of contextual
association in extinction and renewal. 相似文献
9.
Dominic M. Dwyer Hannah L. Pincham Thida Thein Justin A. Harris 《Learning & behavior》2009,37(4):305-310
Learned flavor preferences can be strikingly persistent in the face of behavioral extinction. Harris, Shand, Carroll, and
Westbrook (2004) suggested that this persistence may be due to flavor preference conditioning’s producing a long-lasting change
in the hedonic response to the conditioned stimulus (CS+) flavor. In the present study, the CS+ flavor was presented in simultaneous
compound with 16% sucrose, whereas the CS− flavor was presented with 2% sucrose. During subsequent two- and one-bottle tests,
the CS+ and CS− flavors were presented in 2% sucrose. Hedonic reactions during training and test were assessed using an analysis
of the microstructure of licking behavior. Conditioning resulted in greater consumption of the CS+ than of the CS− that did
not extinguish over repeated two- and one-bottle tests. The mean lick cluster size was higher for the CS+ than for the CS−
only on the first cycle of tests. Since lick cluster size can be used as an index of stimulus palatability, the present results
indicate that although the hedonic reaction to the CSs did change, this was not maintained across repeated tests. Thus, changes
in the hedonic response to the conditioned flavors cannot explain the resistance to the extinction of learned flavor preferences. 相似文献
10.
The present research examined the temporal distribution of responding in a lick suppression paradigm. In Experiment 1, rats were trained with either a 30- or a 120-s conditioned stimulus (CS), which was followed either by a footshock (unconditioned
stimulus [US]) or nothing. Licking during the CS was suppressed only in the former condition. Suppression was more pronounced
early in the CS. In Experiment 2, rats were exposed to two 30-s or two 120-s CSs, with delivery of the shock being contingent on CS1 for half of the animals
and on CS2 for the other half. For both the paired and the unpaired conditions, suppression at the beginning of CS1 was observed
for all the groups. By discounting the possibility of generalization between CS1 and CS2, it appears that this initial suppression
was not a conditioned response to the CS, but an unconditioned one due to mere exposure to the shock US. 相似文献
11.
In four conditioned suppression experiments with rats (Rattus norvegicus), backward pairings of a shock unconditioned stimulus (US) and a tone conditioned stimulus (CS) eliminated an already established conditioned response (CR), but there was recovery of the CR if the shock was later withheld. In Experiment 1, there was recovery after backward pairings, regardless of whether the period after the US was normally shock free or not. In Experiment 2, the occurrence of recovery depended on the CS’s being presented closely after the US in response elimination. Levels of recovery were positively correlated with the resistance of the response to elimination during backward pairings (Experiments 3 and 4). Taken together, these data support the notion that recovery after backward pairings is a form of renewal (see, e.g., Bouton, 1991) and is not due toprotection from extinction. 相似文献
12.
Prior research on Pavlovian-to-instrumental transfer has shown that when a CS previously associated with shock (AvCS+) is presented contingent upon a choice response to a discriminative stimulus for food reinforcement, it facilitates discrimination learning. Conversely, a response-contingent CS previously associated with the absence of shock (AvCS?) retards discrimination learning. To evaluate whether these findings reflect across-reinforcement blocking and enhancement effects, two experiments investigated the effects of appetitively conditioned stimuli on fear conditioning to a novel stimulus that was serially compounded with the appetitive CS during conditioned-emotional-response (CER) training. Although there were no differential effects of the appetitive CSs in CER acquisition, Experiment 1, using a relatively weak shock US, showed that a CS previously associated with food (ApCS+) retarded CER extinction to the novel stimulus, in evidence of enhanced fear conditioning to that stimulus. In addition, Experiment 2, using a stronger shock US, showed that a CS previously associated with the absence of food (ApCS?) facilitated CER extinction to the novel stimulus, in evidence of weaker fear conditioning to that stimulus. These results parallel traditional blocking effects and indicate not only that an ApCS+ and an ApCS? are functionally similar to AvCSs of opposite sign, but that their functional similarity is mediated by common central emotional states. 相似文献
13.
Three experiments demonstrated that, following the extinction of an established conditioned stimulus (CSA—e.g., tone), the
pairing of a novel, cross-modal stimulus (CSB—e.g., light) with the unconditioned stimulus (US) results in strong recovery
of responding to the extinguished CSA. Experiment 1 demonstrated that the recovery of responding to CSA is not the result
of US reinstatement but is attributable to pairings of CSB with the US. Experiment 2 demonstrated that the recovery of responding
is specific to CSA and is not the result of cross-modal generalization. Experiment 3 revealed that a large number of CSB-US
pairings in Stage 1 significantly reduced the amount of recovery to CSA during subsequent CSB-US trials. Experiment 3 also
provided unexpected evidence of cross-modal secondary extinction. The extinction and subsequent recovery of responding seen
in the present experiments is discussed with respect to possible contributions from contextual associations, CS processing,
US processing, conditioned response expression, and layered excitatory associations. 相似文献
14.
In a series of related experiments, we studied associative phenomena in snails (Helix aspersa), using the conditioning procedure of tentacle lowering. Experiments 1A and 1B demonstrated a basic conditioning effect in
which the pairing of an odor (apple) as the conditioned stimulus (CS) with the opportunity to feed on carrot as the unconditioned
stimulus (US) made snails exhibit increased levels of tentacle lowering in the presence of the CS. Experiments 2 and 3 showed
that the magnitude of the conditioning was reduced when snails were exposed to the CS prior to the conditioning trial (a latent
inhibition effect). Experiment 4 examined the effects produced by pairing a compound CS (apple—pear) with food presentations
and demonstrated the existence of an overshadowing effect between the two odors. Experiment 5 revealed that pairing one CS
with another previously conditioned stimulus increased tentacle lowering to the new CS (a second-order conditioning effect).
Finally, Experiment 6 showed that pairing two odors prior to conditioning of one of them promoted an increase in tentacle
lowering in response to the other (a sensory preconditioning effect). The results are discussed in terms of an associative
analysis of conditioning and its implications for the study of cognition in invertebrates. 相似文献
15.
Kenneth A. McNish Stephanie L. Betts Susan E. Brandon Allan R. Wagner 《Learning & behavior》1997,25(1):43-52
Two experiments of Pavlovian conditioning with rabbits evaluated the effects of initiating or continuing a conditioned stimulus (CS) after a paraorbital unconditioned stimulus (US). In Experiment 1, backward pairings, in which a CS came on after the US, produced a CS that appeared inhibitory on a measure of eyeblink conditioning but excitatory on a potentiated-startle measure of conditioned fear. In Experiment 2, extending the duration of a CS that came on prior to the US, so that it continued after the US, decreased eyeblink conditioned responses, whereas it increased conditioned fear. The data from the two experiments confirm and extend those of Tait and Saladin (1986), supporting the suppositions of AESOP (Wagner & Brandon, 1989) that conditioned eyeblink and conditioned fear can be dissociated under various temporal relationships between the CS and US. 相似文献
16.
In two experiments, we examined the conditions under which signaling an unconditioned stimulus (US) with a nominal conditioned stimulus (CS) interferes with the conditioning of situational cues in defensive freezing in the rat. Subjects received footshock USs that were (1) either signaled or unsignaled and (2) either varied or fixed in their temporal location within the conditioning session. Experiment 1, with only one trial per session, yielded no evidence that signaling affected pretrial freezing using either a fixed or variable interval between placement in the context and shock onset. In a test in which no CSs or footshocks were presented, groups that previously had received footshock at a fixed temporal location showed greatest freezing at around that same time. For groups that had received footshocks at various times, freezing declined across the test session. Experiment 2 showed overshadowing of pretrial freezing after more extensive conditioning with many trials per session, but only if the intershock intervals were variable rather than fixed. 相似文献
17.
Todd R. Schachtman Wesley J. Kasprow Robert C. Meyer Mark J. Bourne Julie A. Hart 《Learning & behavior》1992,20(3):207-218
Using a conditioned taste aversion preparation overshadowing of flavor-illness association was produced through the presentation of a second flavor during the interval between the first flavor and illness. The modulatory effects of extinguishing the association between the second (over-shadowing) flavor and illness on conditioned responding to the target flavor was investigated. In Experiment 1, we found that, following one-trial overshadowing, extinction of the overshadowing flavor had no effect on conditioned responding to the target flavor. In Experiment 2, we found a similar absence of an effect of extinction of the overshadowing stimulus in a multitrial over-shadowing paradigm. Experiment 3 confirmed the results of Experiments 1 and 2 using conditioning parameters that were designed to weaken the association between the overshadowed flavor and illness. In Experiments 4 and 5, we used simultaneous presentation of the flavors during conditioning and obtained a weakened aversion to the overshadowed flavor when the overshadowing CS was extinguished. These findings are inconsistent with previous observations in conditioned fear preparations that suggest that extinction of the association between the overshadowing stimulus and the unconditioned stimulus attenuates overshadowing. Possible reasons for the discrepant results are discussed. 相似文献
18.
Conditioned lick suppression in rats was used to explore the role of timing in trace conditioning. In Experiment 1, two groups of rats were exposed to pairings of a CS (CS1) with a US, under conditions in which the interstimulus interval (ISI) that separated CS1 offset and US onset was either 0 or 5 sec. Two additional groups were also exposed to the same CS1→US pairings with either a 0 or a 5-sec ISI, and then received “backward” second-order conditioning in which CS1 was immediately followed by a novel CS2 (i.e., CS1→CS2). A trace conditioning deficit was observed in that the CS1 conditioned with the 5-sec gap supported less excitatory responding than the CS1 conditioned with the 0-sec gap. However, CS2 elicited more conditioned responding in the group trained with the 5-sec CS1-US gap than in the group trained with the 0-sec CS1-US gap. Thus, the CS1-US interval had inverse effects on first- and second-order conditioned responding. Experiment 2 was conducted as a sensory preconditioning analogue to Experiment 1. In Experiment 2, rats received the CS1?CS2 pairings prior to the CS1→US pairings (in which CS1 was again conditioned with either a 0 or a 5-sec ISI). Experiment 2 showed a dissociation between first- and second-order conditioned responding similar to that observed in Experiment 1. These outcomes are not compatible with the view that differences in responding to CSs conditioned with different ISIs are mediated exclusively by differences in associative value. The results are discussed in the framework of the temporal coding hypothesis, according to which temporal relationships between events are encoded in elementary associations. 相似文献
19.
Peter C. Holland 《Learning & behavior》1979,7(4):424-432
The form of rats’ Pavlovian conditioned responses to visual and auditory conditioned stimuli (CSs) paired with a variety of unconditioned stimuli (USs) was examined in three experiments using direct behavioral observation techniques. In Experiment 1, the form of conditioned behavior occurring most frequently during later portions of the CS-US interval depended only on which of several appetitive USs was used, but the form of behavior occurring most frequently during early portions of the CS-US interval depended only on the nature of the CS. US-dependent behaviors resembled the response to the US, and CS-dependent behaviors resembled the original orienting response (OR) to the CS. In Experiment 2, the use of larger magnitude appetitive USs resulted in higher frequencies of US-dependent behaviors, but lower frequencies of CS-dependent behaviors in the presence of auditory and visual CSs. In Experiment 3, US-dependent conditioned behavior to auditory and visual CSs paired with shock was more frequent when high-intensity shocks were used, but CS-dependent behavior was more frequent when low-intensity shocks were used. These results suggested that Pavlovian conditioned responding may involve two independent types of behavior—one appropriate to the US and another based on the original OR to the CS. 相似文献
20.
Dale Swartzentruber 《Learning & behavior》1993,21(1):14-22
Two autoshaping experiments with pigeons examined contextual control of responding to conditioned stimuli (CSs) when separate phases of reinforcement and nonreinforcement occurred in different contexts. In Experiment 1, a CS was first conditioned in Context A prior to nonreinforcement in Context B. In Experiment 2, a conditional discrimination reversal procedure was employed in which one CS was first reinforced in A prior to nonreinforcement in B, and another CS was first nonreinforced in A and then reinforced in B. In both experiments, the extent of contextual control was assessed by testing the CSs in A and B. The CS specificity of control was assessed by examining control of CSs that had been treated like the original CSs, but in a different pair of contexts. The results show that contexts control responding to CSs through a CS-specific mechanism, as well as through a mechanism that is independent of the identity of the CS. However, the extent to which control is mediated by a CS-independent mechanism depends on the training history of the CS. 相似文献