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1.
The present paper is an attempt to throw some light on two Composite genera,
Cavea W. W. Smith et Small and Nannoglottis Maxim., of which the generic features
and the systematic position were heretofore rather badly known.
Cavea, a monotypic genus based upon Saussurea salwinensis Drumm., was described
by W. W. Smith and Small as monocious. From the ample material now in our com-
mand, it is found that the genus is either monocious and dioecious, the capitula in the
latter case being homogamous. The individuals exclusively provided with sterile flowers
are usually smaller in size, sometimes even stemless. Such repartition of sexes was
known in the subtribe Plucheinae (tribe Inuleae), to which W. W. Smith and Small were
correct to ascribe their genus.
Nannoglottis was considered by Maximovicz as un abnormal genus of the tribe
Inuleae, having heterogamous capitula with one series of peripheric fertile ligulate pistil-
late flowers. Having examined the type specimen of N. carpesioices Maxim. and the ex-
tensive collection from Kansu and Tsinghai, we incline to say that the genetic characters
as discerned by Maximovicz are doubtlessly due to his inaccurate observation. The
flowers of Nannoglottis are in fact trimorphous, of which the pistillate ones are 2-3-
seriate, the outer being shortly ligulate and the inner, short-tubulate with truncate apex.
Handel-Mazzetti, much later, indicated that, in another species, the pistillate flowers are
2-3-seriate, but no mention was made about the inner tubulate pistillate flowers.
Franchet, appearing to be unware of Maximovicz's genus, founded his Stereosanthus
Franch. and considered it as a genus intermediate between the tribes Inuleae and Sene-
cioneae. The genus was misinterpreted by Franchet as having dimorphous flowers, the
pistillate ones being all ligulate. Curious enough, one of the major generic features, viz.,
the presence of inner tubulate pistillate flowers, was neglected by all early botanists and
was first recognized by Handel-Mazzetti fourty years later.
However, Handel-Mazzetti was still unacquainted with the characters of Nannoglot-
tis and proposed therefore his new genus Vierhapperia Hand.-Mzt. on the reason that the
flowers are trimorphous. Moreover, he compared his genus with Conyza L. and Erigeron
L. of the tribe Astereae, but the characters marked by him appear not to be different
from those of Nannoglottis.
From the above morphological comparaison, it is evident that Nannoglottis and two
other genera in question possess in common important characters in the general appea-
rance, the flower-forms and the structure of achenes and pappus. The differences in the
disposition of involucral bracts and the length of ligules are criteria good for separating
species, but can hardly be regarded as sufficient for generic delimitation. Moreover,
these three genera are almost similar in their geographical distribution. These points are
adequate to warrant that Stereosanthus and Vierhapperia are in reality congeneric with
Nannoglottis.
Nannoglottis appears to have been correctly placed by Hoffmann in the subtribe
Senecioninae (tribe Senecioneae) though in that subtribe it finds no close relatives. On
account of the outstanding characters of the genus, a further study of its systematic posi-
tion is required.
The present paper is brought to close by a tentative scheme of classification of the
genus so amplified, of which two sections, namely, sect. Stenolepis Ling et Y. L. Chen
(Stereosanthus Franch., p. p. min.) and sect. Nannoglottis (Stereosanthus Franch., p. p.maxim.; Vierhapperia Hand.-Mzt.), altogether including 9 species, are being proposed. 相似文献
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5.
石铸 《中国科学院研究生院学报》1987,25(3):189-203
After having examined all specimens of the genus Prenanthes L. of Compositae in
the Herbarium of Institute of Botany, Academia Sinica, I find that the classic concept on the
genus Prenanthes established by G. Bentham in 1873 has not been held exactly by some of Euro-
pean, American, Japanese and Chinese botanists. For example, W. B. Hemsley, S. T. Dunn, A.
Franchet, S. Kitamura and C. C. Chang placed plants from China which belong to other groups
into the genus; I also find that the classic concept of the genus is not clear. The present paper
makes a revision not only on the classic concept of the genus, but also on its concept assumed by
the above-mentionded botanists.
With the combination of numerous (25-35), white or yellow ligular florets, numerous ribs
of achenes, Prenanthes alba L. (Nabalus albus (L.) Shih, comb. nov.) is distinctly different
from Prenanthes purpurea L., which has the combination of purple, few (5-15) ligular flo-
rests and few ribs of achenes. Nabalus Cass., as a genus established early (1825) by H. Cassini,
should be restored. It is not reasonable to treat Nabalus as a subgenus (E. B. Babcock et al.
1947) or a section (S. Kitamura, 1956) or as a synonym (G. Bentham, 1983) of the genus
Preanathes L.
The present author recognizes seven species in the new revised genus Prenanthes L. in
China, 4 of which are described as new. In the genus Nabalus Cass. only one species, N.
ochroleuca Maxim., is distributed in Northeast China.
As Lactuca melanantha Franch. (1895), Prenanthes henryi Dunn (1903), P. glandulosa
Dunn (1903), Lactuca triflora Hemsl. (1888) (it was transferred to Prenanthes L. by C. C.
Chang in 1934), Prenanthes formosana Kitam. (1934) and P. wilsoni Chang (1934) all have
campanulate involucres, purple phyllaries, purple dorsi-ventrally compressed achenes, longitu-
dinal rids 6-9 on each side of achene truncate and beakless at its apex and pilose tubes of co-
rollae, they should be placed neither into the genus Prenanthes with obtusely tri-or pentagonous,
subterete achenes and glabrous tubes of corollae, nor into the genus Lactuca with beak achenes.
Besides the above-mentioned species misnamed by some of foreign and Chinese botanists, 6 other
species also have the same structure in achenes and corollae. Evidently, they fall into a new
genus with the name Notoseris Shih.
The new genus Notoseris Shih of the tribe Lactuceae of Compositae seems to be more re-
miniscent of Lactuca L. than of Prenanthes L. emend.
All the 12 species of the genus Notoseris Shih are endemic to China and distributed in the
area of south of Yantze River. Of them 6 are new combinations and 6 are described as new. 相似文献
6.
在比较研究标本室标本,包括模式标本以及野外考察的基础上,确认Chaetoseris hispida Shih 与Ch. cyanea (D. Don) Shih为同种植物,并把前者作为后者的异名处理。 相似文献
7.
按照花粉外壁表面纹饰的不同,可将春黄菊族中下列各属分为两大类群:(1)明显具刺:亚菊
属Ajania Poljak., 女蒿属Hippolytia Poljak., 百花蒿属Stilpnolepis Krasch.。 (2)具退化状小刺:紊蒿属
Elachanthemum Ling et Y.R.Ling,线叶菊属Filifolium Kitam., 画笔菊属Ajaniopsis Shih,喀什菊属
Kaschgaria Poljak.),栉叶蒿属Neopallasia Poljak.,绢蒿属Seriphidium Poljak.。根据花粉形态特征,我们认为将紊蒿属和百花蒿属仍分为两个相互独立的属较为适宜。 相似文献
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近日.与美国哈佛大学植物标本馆David E.Boufford博士合作.通过多重证据成立了喜马拉雅-横断山区特有属:假合头菊属。 相似文献
10.
张芝玉 《中国科学院研究生院学报》1982,20(4):402-409
1. The present paper describes the observations of chromosome numbers and
karyomorphology of 2 species of 2 endemic genera and I endemic species of Chinese
Ranunculaceae: Asteropyrum peltatum (Franch.) Drumm et Hutch. 2n=16, x=8;
Kingdonia unifolia Balf. f. et W. W. Sm. 2n=18, x=9 and Calathodes oxycarpa Spra-
gue 2n=16, x=8. The chromosome counts of three ranunculaceous genera are repor-
ted for the first time.
2. The morphylogical, palynological and cytological date in relation to the syste-
matic postition of Asteropyrum, Kingdonia and Calathodes within the family Ranun-
culaceae are diseussed and resulted in following conclusions:
(1). On the basis of the basic number x=8 in Asteropyrum, it is further con-
firmed that this genus is distinct from the r elated genera such as Isopyrum, Dichocarp-
um and other allied taxa.
The comparison of Asteropyrum with Coptis shows that they are identical in short
chromosomes, with magnoflorina and benzylisaquinodine type of alkaloides, but dif-
ferent from coptis in the chromosome numbers (T-type), pantocolpate pollens, united
carpels and the dorsi-ventral type of petioles. In view of these fundamental morpho-
logical and cytological differences, Asterop yrum is better raised to the level of Tribe.
However Asteropyrum and Coptis may represent two divaricate evolutional lines of
Thalictroideae.
(2). The systematic position of the genus Kingdonia has been much disputed in
the past. We support the view of Sinnote (1914), namely, the trilacunar in leaf
traces “the ancient type”, appeared in the angiosperm line very early, while the uni-
lacunar of Kingdonia may be derived from the trilacunar. On the basis of the chromo-
some numbers and morphylogical observation, the present writer accept Tamura’s and
Wang’s treatment by keeping Kingdonia in Ranunculaceae instead of raising it to a
family rank as has been been done by Forster (1961). Kingdonia and Coptis are
similar in having short chromosome with x=9, but with one-seeded fruits; therefore it
is suggested that placed into Thalictroideae as an independent tribe, indicating its close
relationship with Coptideae.
(3). Comparing with its allies, Calathodes being with out petals, seems to be more
primitive than Trollius. But Calathodes differs from Trollius with R-type chromosomes in having T-type chromosome with x=8 and subterminal centromere. Those charac-
teristics show that it is very similar to the related genera of Thalictroideae. But as
Kurita already pointed out that most speci es of Ranunculus have usually large long
chromosomes but some species have compar ativelly short chromosomes, therefore we
regard T-type and R-type chromosomes appear independently in different subfamilies
of Ranunculaceae. According to Tamura, G alathodes seems to be closely related to
Megaleranthis, because of the resemblance in follicles. But due to lack of cytological
data of the latter genus, the relationship between the two genera still is not clear pen-
ding further studies. From the fact that the morphology and chromosomes of the
Calathodes differs from that of all other genera of the Helleboroideae, we consider
Calathodes may form an independent tribe of its own with a closer relationship withTrollieae. 相似文献
11.
石铸 《中国科学院研究生院学报》1986,24(4):292-296
Almost no differences are seen in habit and infrescence between the gene-
ra Vladimiria and Dolomiaea; the former that was founded on the basis of Jurinea sal-
winensis Hand.-Mazz. by Iljin should be transferred to the later that was earlier found-
ed on the basis of D. macrocephala DC. by De Candolle. But considering the style which
has fine, spreading and apex-acute branches in the group Vladimiria, it is reasonable
to treat the group Vladimiria as a section in the genus Dolomiaea, i.e. Dolomiaea sect.
Vladimiria (Iljin) Shih. Therefore, the genus Dolomiaea is expanded in the circums-
cription in the present paper, not only including the primary group with short, com-
pact and apex-rounded style branches, i.e. Dolomiaea sect. Dolomiaea, but also sect. Vla-
dimiria (Iljin) shih with different characters of style-arms.
Nine species and 1 variety as new combinations are listed in the sect. Vladimiria
(Iljin) Shih as following: D. denticulata (Ling) Shih, D. forrestii (Diels) Shih, D.
scabrida (Shih et S. Y. Jin) Shih, D. souliei(Franch.) Shih, D. souliei (Franch) Shih
var. mirabilis (Anth.) Shih, D. berardioidea (Franch.) Shih, D. georgii (Anth.) Shih,
D. edulis (Franch.) Shih, D. salwinensis (Hand.-Mazz.) Shih and D. platylepis (Hand.
-Mazz.) Shih. 相似文献
12.
13.
石铸 《中国科学院研究生院学报》1988,26(6):418-428
Having mixed extremely various patterns of achenes for a long time in
the genus Lactuca L., especially showed in the classification of it on the mainland of Asia,
Lactuca L., the primary unnatural genus, becomes more nuclear in its limit. The present
paper makes a revision to attempt providing a clear cline between Lactuca L. and its relative
genera.
On my opinion, only plants, represented by Lactuca sativa L., which have ovoid capitu-
lum during its fruiting, numerous (7-25), yellow ligular florets and longitudinal 1-10 ribs
or striae on each side of achenes, acute into filiform beak at its apex, should be defined as
Lactuca L.
In view of the present concept of Lactuca L., another group of plants, embodied by
Lactuca indica L., having its broadening, thin winged-margin of, 1-3 striae on each side of
achenes, black, dorsi-ventral compressed, acute into thick and short beak at its apex, evident-
ly, should not be placed into Lactuca L., but be regarded as genus, i.e. Pterocypsela Shih,
gen. nov. Pterocypsela Shih distributes in Eastern and southern Asia, where Lactuca L. does
not occur. In China there is all of species, seven species, of Pterocypsela Shih. They are P.
raddeana (Maxim.) Shih, P. indica (L.) Shih, P. elata (Hemsl.) Shih P. lacciniata (Houtt.)
Shih, P. triangulata (Maxim.) Shih, P. sonchus (Lévl.) Shih and P. formosana (Maxim.)
Shih, But in Mediterranean regions and Middle and Western Asia mainly distributes Lactu-
ca L., in which there are only seven species from western China, Xijiang autonomic district.
They are L. dolichophylla Kitam., L. sativa L., L. altaica Fisch. et Mey., L. serriola Torner,
L. dissecta Don, L. auriculata DC. and L. undulata Ledb.
With the combination of violet-purple ligular florets non-ovoid capitulum during its
fruiting, a little thick achenes, acuminate into longer. or short beak at its apex and 4-6 ribs
on each side of it, Mulgedium tataricum (L.) DC. is evidently different from Lactuca sativa L.
with combination of ovoid capitulum during its fruiting, yellow ligular florets and compres-
sed achenes, acute into filiform beak at its apex. There seems to be no point in refusing Mu-
lgedium Cass. as a genus. Strictly speak, Mulgedium Cass. seems to be reminiscent of Para-
prenanthes Chang (see bellow) without beak at the apex of its achenes. In Hengduan moun-
tains and mountain range of Himalayas M. tataricum (L.) DC. discoveries its relative par-
tners. They are M. bracteatum (Mook. f. ex. C. B. Clarke) Shih, M. lessertianum (Wall. ex
C. B. Clarke) DC., M. monocephalum (Chang) Shih, M. umbrosum (Dunn) Shih, M. meridi-
onale Shih and M. polypodifolium (Franch.) Shih.
Having broadering and thickening margin and 4-7 striae on each side of achenes, grey,
dorsi-ventral compressed, truncate and beakless at its apex, Lactuca sibirica (L.) Benth. ex
Maxim. (Sonchus sibiricus L.) is not only different from Lactuca sativa L. with the combi-
nation of yellow ligular florets and filiform beak at apex of its achenes, but also from the
genus Mulgedium Cass. with beak at apex of its achenes. Therefore, Lagedium Sojak, establi-
shed by J. Sojak (1961), should be restored. It should be point out that Lagedium Sojak is
monotypic genus excluding Mulgedium tataricum (L.) DC. with beak at the apex of its ache-
nes, non-marginated. Lagedium Sojak widely distributes in temperate and frigid zone of
northern hemisphere. In our country, Lagedium sibiricum (L.) Sojak, sole species of the ge-
nus, restricts its northeast region.
Paraprenanthes Chang, the new genus, established by Ch. Ch. Chang (1950) based on La-
ctuca sororia Miq. and Lactuca yunnanensis Franch. seems to be a distinct genus. This ge-
nus is characterized by 4-6 ribs on each side of achenes, black, fusiform, a little thick and
beakless at its apex. Besides above-mentioned two species, assigned by Chang, in forest of tro-
pic and subtropic zones on the mainland of Asia there are many species with same structure
in achenes like L. sororia Miq. and L. yunnanensis Franch. The genus Paraprenanthes Chang
seems to be reminiscants of Mulgedium Cass. and Lagedium Sojak, but differs from the former
in its beakless achenes, from the later in its non-marginated achenes and 4-6 ribs on each
side of it. In our country, there are all species of the genus known in the tropic and sub-
tropic zones on the mainland of Asia. They are P. sororia (Miq.) Shih, P. yunnanensis (Fra-
nch.) Shih, P. longiloba Ling et Shih, sp. nov., P. heptontha Shih et D. J. Liou, sp. nov., P.
prenanthoides (Hemsl.) Shih, P. pilipes (Migo) Shih, P. thirionii (Lévl.) Shih, P. sylvicola
Shih., sp. nov., P. multiformis Shih, sp. nov., P. auriculiformis Shih, sp. nov. and P. sagitti-
formis Shih, sp. nov. 相似文献
14.
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石铸 《中国科学院研究生院学报》1993,31(6):533-548
本文采用狭义概念,将苦荬菜类群(Ixeris group)处理为4个属,即苦荬菜属Ixeris Cass.沙苦荬属Chorisis DC.小苦荬属Ixeridium(A.Gray)Tzvel.及黄瓜菜属Paraixeris Nakai并提供了分属、种检索表。文中记载了3个新种及7个新组合种。 相似文献
16.
17.
石铸 《中国科学院研究生院学报》1991,29(5):394-417
本文是对中国-喜马拉雅地区的原莴苣属植物提出的分类订正。是笔者前一篇文章《莴苣属订正
及亚洲大陆菊科植物二新属》的续篇。文中建立了两个新属,即毛鳞菊属Chaetoseris Shih和细莴苣属Stenoseris Shih;文末及文中提供了本文所涉及的一些属种的分属和分种检索表。 相似文献
18.
风毛菊属Saussurea DC.是菊科物种分化十分剧烈和分类处理十分困难的一个属。该属的单系起源性质、属下分类系统以及一些独特形态物种的系统位置尚不清楚,有待进一步验证。本文测定了代表该属5个亚属37种植物43个样品和川木香属Dolomiaea DC.的1种样品的叶绿体DNA trnL-F序列,并调取菜蓟族Cardueae Cass.与风毛菊属具有一定亲缘关系的13属的该序列,一起进行了分支分析,重点验证该属的属下形态分类系统以及形态特殊、青藏高原地区特有的雪兔子亚属subgen. Eriocoryne 相似文献
19.
铁线莲属研究随记(Ⅲ) 总被引:2,自引:0,他引:2
王文采 《中国科学院研究生院学报》2000,38(6):497-514
(1)在铁线莲属的15个组330余种中,只有绣球藤组sect.Cheiropsis的5个种(短梗铁线莲Clematis brevipes Rehd.,美花铁线莲C.potaninii Maxim.,绣球藤C.montana Buch.-Ham.ex DC.,薄叶铁线莲C.gracilifolia Rehd.&; Wils.和金毛铁线莲C.chrysocoma Franch.)的花与比自己原始的近缘属——银莲花属Anemone的花在构造上相近似,这表现在:萼片水平开展,膜质,倒卵形,顶端圆形或圆钝,内面无毛,外面被柔毛,但边缘无狭的短绒毛带;雄蕊无毛,花丝条形(linear)或狭条形,花药长圆形或狭长圆形,药隔顶端不突出,上述均是铁线莲属花的原始特征。在此属的其他种,可以看到萼片和雄蕊的形态发生了以下诸种变化:萼片从平展变为斜上方开展或甚至直立,质地变厚,呈纸质,有时甚至呈革质,形状多少变狭长,呈倒卵状长圆形、长圆形、倒披针形、披针形、条形,甚至狭条形,顶端有时变为渐尖或渐狭,内面出现了或疏或密的柔毛,外面边缘出现了一条极狭的短绒毛带;在雄蕊方面,花丝全长由等宽变为不等宽,呈狭披针形或狭倒披针形,花药变狭长,呈条形或狭条形,药隔顶端出现了短或长的突起,花丝或花药或二者上出现了或多或少的柔毛。上述的形态变化当均属衍生特征;(2)建立了绣球藤组sect.Cheiropsis和广布于北、南美洲和大洋洲具单性花的单性铁线莲组Sect.Aspidanthera的新分类系统;(3)描述了1新亚组,13新系,5新种,4新变种;(4)做出了6新等级,2新组合。 相似文献