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1.
木兰科分类系统的初步研究 总被引：10，自引：0，他引：10
刘玉壶 《中国科学院研究生院学报》1984,22(2):89-109
A new system of classification of Magnoliaceae proposed. This paper deals mainly with taxonomy and phytogeography of the family Magnoliaceae on the basis of external morphology, wood anatomy and palynology. Different authors have had different ideas about the delimitation of genera of this family, their controversy being carried on through more than one hundred years (Table I). Since I have been engaged
in the work of the Flora Reipublicae Popularis Sinicae, I have accumulated a considerable amount of information and material and have investigated the living plants at their natural localities, which enable me to find out the evolutionary tendencies and primitive morphological characters of various genera of the family. According to the evolutionary tendencies of the characters and the geographical distribution of this family I propose a
new system by dividing it into two subfamilies, Magnolioideae and Liriodendroideae Law (1979), two tribes, Magnolieae and Michelieae Law, four subtribes, Manglietiinae Law, Magnoliinae, Elmerrilliinae Law and Micheliinae, and fifteen genera (Fig. 1 ), a system which is different from those by J. D. Dandy (1964-1974) and the other authors.
The recent distribution and possible survival centre of Magnoliaceae. The members of Magnoliaceae are distributed chiefly in temperate and tropical zones of the Northern Hemisphere, ——Southeast Asia and southeast North America, but a few genera and species also occur in the Malay Archipelago and Brazil of the Southern Hemisphere. Forty species of 4 genera occur in America, among which one genus (Dugendiodendron) is endemic to the continent, while about 200 species of 14 genera occur in Southeast Asia, of which 12 genera are endemic. In China there are about 110 species of 11 genera which mostly occur in Guangxi, Guangdong and Yunnan; 58 species and more than 9 genera occur in the mountainous districts of Yunnan. Moreover, one genus
(Manglietiastrum Law, 1979) and 19 species are endemic to this region. The family in discussion is much limited to or interruptedly distributed in the mountainous regions of Guangxi, Guangdong and Yunnan. The regions are found to have a great abundance of species, and the members of the relatively primitive taxa are also much more there than in the other regions of the world.
The major genera, Manglietia, Magnolia and Michelia, possess 160 out of a total of 240 species in the whole family. Talauma has 40 species, while the other eleven genera each contain only 2 to 7 species, even with one monotypic genus. These three major genera are sufficient for indicating the evolutionary tendency and geographical distribution of Magnoliaceae. It is worthwhile discussing their morphological characters and
distributional patterns as follows:
The members of Manglietia are all evergreen trees, with flowers terminal, anthers dehiscing introrsely, filaments very short and flat, ovules 4 or more per carpel. This is considered as the most primitive genus in subtribe Manglietiinae. Eighteen out of a total of 35 species of the genus are distributed in the western, southwest to southeast Yunnan. Very primitive species, such as Manglietia hookeri, M. insignis and M. mega-
phylla, M. grandis, also occur in this region. They are distributed from Yunnan eastwards to Zhejiang and Fujian through central China, south China, with only one species (Manglietia microtricha) of the genus westwards to Xizang. There are several species distributing southwards from northeast India to the Malay Archipelago (Fig. 7).
The members of Magnolia are evergreen and deciduous trees or shrubs, with flowers terminal, anthers dehiscing introrsely or laterally, ovules 2 per carpel, stipule adnate to the petiole. The genus Magnolia is the most primitive in the subtribe Magnoliinae and is the largest genus of the family Magnoliaceae. Its deciduous species are distributed from Yunnan north-eastwards to Korea and Japan (Kurile N. 46’) through Central
China, North China and westwards to Burma, the eastern Himalayas and northeast
India. The evergreen species are distributed from northeast Yunnan (China) to the
Malay Archipelago. In China there are 23 species, of which 15 seem to be very primi-
tive, e.g. Magnolia henryi, M. delavayi, M. officinalis and M. rostrata, which occur in
Guangxi, Guangdong and Yunnan.
The members of Michelia are evergreen trees or shrubs, with flowers axillary, an-
thers dehiscing laterally or sublaterally, gynoecium stipitate, carpels numerous or few.
Michelia is considered to be the most primitive in the subtribe Micheliinae, and is to
the second largest genus of the family. About 23 out of a total of 50 species of this
genus are very primitive, e.g. Michelia sphaerantha, M. lacei, M. champaca, and M.
flavidiflora, which occur in Guangdong, Guangxi and Yunnan (the distributional center
of the family under discussion) and extend eastwards to Taiwan of China, southern
Japan through central China, southwards to the Malay Archipelago through Indo-China.
westwards to Xizang of China, and south-westwards to India and Sri Lanka (Fig. 7).
The members of Magnoliaceae are concentrated in Guangxi, Guangdong and Yunnan
and radiate from there. The farther away from the centre, the less members we are
able to find, but the more advanced they are in morphology. In this old geographical
centre there are more primitive species, more endemics and more monotypic genera.
Thus it is reasonable to assume that the region of Guangxi, Guangdong and Yunnan,
China, is not only the centre of recent distribution, but also the chief survival centreof Magnoliaceae in the world. 相似文献
2.
张志耘 《中国科学院研究生院学报》1988,26(5):394-403
The morphological characters in the genus Orobanche were evaluated from the
taxonomic point of view. The author finds that the plants of this genus are relatively similar
to each other in respect to characters of vegetative organs, fruits and seeds. But the differences
in the floral structures can be served as a basis for delimitating infrageneric taxa. The seed
coat of 18 species and pollen grains of 6 species were also examined under scanning electron
microscope (SEM). They seem to have little significance for distinguishing species.
The result supports G. Beck’s (1930) division of the genus Orobanche into 4 sections, of
which 2 occur in China, based on the characters of the inflorescence, bracteoles and calyx.
The author considers that some characters, such as anther hairy or not, upper lip of corolla
entire or not, lower lip longer or shorter than the upper one, the state of corolla-tube inflec-
tion and the hair type of filaments and plants, are important in distinguishing Chinese species.
A key to the species of Orobanche in China is given.
This genus consists of about 100 species, and is mostly confined to Eurasia, with over 60
species found in Caucasus and Middle Asia of USSR, where may be the mordern distribu-
tional centre.
Orobanche L. in China is represented by 23 species, 3 varieties and l forma. As shown in
Table 1, most species (12 species) are found in Xinjiang, which clearly shows a close floristic
relationship between this region and Middle Asia of USSR. 6 species are endemic to China,
of which 4 are confined to the Hengduan Mountains (Yangtze-Mekong-Salwin divide).
The relationships between this genus and related ones of Orobanchaceae are also discussed.
The author holds the following opinions: the genus Phelypaea Desf. should be considered as a
member of Orobanche L. Sect. Gymnocaulis G. Beck, the monotypic genus, Necranthus A.
Gilli endemic to Turkey, is allied with Orobanche L. Sect. Orobanche, the monotypic genus,
Platypholis Maxim, endemic to Bonin Is. of Japan, is far from Orobanche L. in relation and
should be regarded as a separate genus.
The 11 OTU’s, including all the sections of Orobanche L. and 7 genera of Orobanchaceae,
and 15 morphological characters were used in the numerical taxonomic treatment to test the
above-mentioned suggestions. After standardization of characters, the correlation matrices were
computerized. The correlation matrices were made to test the various clustering methods. At
last the UPGMA clustering method was chosen and its result is shown in a phenogram. The
result of numerical analysis is basically in accordance with the suggestions. 相似文献
3.
Xizang (Tibet) is rich in Leguminosae flora, comprising 41 genera and 254 species
so far known, exclusive of the commonly cultivated taxa (including 11 genera and 16
species). There are 4 endemic genera (with 8 species), 10 temperate genera (with 175
species) and 19 tropical genera (with 46 species) as well as the representatives of those
genera whose distribution centers are in East Asia-North America, Mediterranean
and Central Asia.
1. There are altogether 4 endemic genera of Leguminosae in this region. Accord-
ing to their morphological characters, systematic position and geographical distribution,
it would appear that Salweenia and Piptanthus are Tertiary paleo-endemics, while
Straceya and Cochlianths are neo-endemics. Salweenia and Piptanthus may be some
of more primitive members in the subfamily Papilionasae and their allies are largely
distributed in the southern Hemisphere. The other two genera might have been derived
from the northern temperate genus Hedysarum and the East Asian-North American
genus Apios respectively, because of their morphological resemblance. They probably
came into existanc during the uplifting of the Himalayas.
2. An analysis of temperate genera
There are twelve temperate genera of Leguminosae in the region, of which the
more important elements in composition of flora, is Astragalus, Oxytropis and Cara-
gana.
Astragalus is a cosmopolitan genus comprising 2000 species, with its center
distribution in Central Asia. 250 species, are from China so far known, in alpine zone of
Southwest and Northwest, with 70 species extending farther to the Himalayas and
Xizang Plateau.
Among them, there are 7 species (10%) common to Central Asia, 12 species (15.7%)
to Southwest China and 40 species (60%) are endemic, it indicates that the differentia-
tion of the species of the genus in the region is very active, especially in the subgenus
Pogonophace with beards in stigma. 27 species amounting to 78.5% of the total species of
the subgenus, are distributed in this region. The species in the region mainly occur in
alpine zone between altitude of 3500—300 m. above sea-level. They have developed into
a member of representative of arid and cold alpine regions.
The endemic species of Astragalus in Xizang might be formed by specialization of
the alien and native elements. It will be proved by a series of horizontal and vertical vicarism of endemic species. For example, Astragalus bomiensis and A. englerianus are
horizontal and vertical vicarism species, the former being distributed in southeast part
of Xizang and the latter in Yunnan; also A. arnoldii and A. chomutovii, the former
being an endemic on Xizang Plateau and latter in Central Asia.
The genus Oxytropis comprises 300 species which are mainly distributed in the
north temperate zone. About 100 species are from China so far known, with 40 species
extending to Himalayas and Xizang Plateau. The distribution, formation and differ-
entiation of the genus in this region are resembled to Astragalus. These two genera are
usually growing together, composing the main accompanying elements of alpine mea-
dow and steppe.
Caragana is an endemic genus in Eurasian temperate zone and one of constructive
elements of alpine bush-wood. About 100 species are from China, with 16 species in Xi-
zang. According to the elements of composition, 4 species are common to Inner Mon-
golia and Kausu, 4 species to Southwest of China, the others are endemic. This not only
indicates that the species of Caragana in Xizang is closely related to those species of
above mentioned regions, but the differentiation of the genus in the region is obviously
effected by the uplifting of Himalayas, thus leading to the formations of endemic species
reaching up to 50%.
3. An Analysis of Tropical Genera
There are 19 tropical genera in the region. They concentrate in southeast of Xizang
and southern flank of the Himalayas. All of them but Indigofera and Desmodium are
represented by a few species, especially the endemic species. Thus, it can be seen that
they are less differentiated than the temperate genera.
However, the genus Desmodium which extends from tropical southeast and northeast
Asia to Mexio is more active in differentiation than the other genera. According to Oha-
Shi,s system about the genus in 1973, the species of Desmodium distributed in Sino-Hima-
laya region mostly belong to the subgenus Dollinera and subgenus Podocarpium. The
subgenus Dollinera concentrates in both Sino-Himalaya region and Indo-China with 14
species, of which 7 species are endemic in Sino-Himalaya. They are closely related to
species of Indo-China, southern Yunnan and Assam and shows tha tthey have close con-
nections in origin and that the former might be derived from the latter.
Another subgenus extending from subtropical to temperate zone is Podocarpium.
Five out of the total eight species belonging to the subgenus are distributed in Sino-
Himalaya and three of them are endemic.
An investigation on interspecific evolutionary relationship and geographic distribu-
tion of the subgenus shows that the primary center of differentiation of Podocarpium
is in the Sino-Himalaya region.
Finally, our survey shows that owing to the uplifting of the Himalayas which has
brought about complicated geographic and climatic situations, the favorable conditions
have been provided not only for the formation of the species but also for the genus in cer-tain degree. 相似文献
4.
张若蕙 《中国科学院研究生院学报》1984,22(2):110-118
The first classification for the genus Ormosia was proposed by Bentham. It was
followed by Taubert (1892) in Engler and Prantl’s Nat. Pflanzenf., who divided the genus
into 2 sections. On the basis of the pod structure and seed characters Prain (1900) arran-
ged the genus in 2 sections with 4 subsections. In the monograph on the genus Merrill
and L. Chen ( 1943 ) limited their taxonomic study to Chinese and Indo-Chinese species, and
recognized 34 species and 15 series. Recently Yakovlev (1971-1976) has treated the ge-
nus in 6 separate genera.
In the present paper the author recognizes 35 species, of which 7 species and 2 varie-
ties are new. The Chinese species of the genus are grouped into 3 sections and 6 series inmy classification. 相似文献
5.
王文采 《中国科学院研究生院学报》1980,18(3):266-282
1. Having analyzed the external morphology of the genus Microula, the author
has proposed a series of criteria as bases for the construction of a classification scheme
of this genus. The most important ones are as follows:
1) The normally developed stem is primitive, and the strongly abbreviated stem
more advanced.
2) The small inconspicuous bracts are more primitive than the large suborbicular
densely arranged ones, which almost entirely cover the flowers and the fruits.
3) Nutlets with small dorsal pit are more primitive than those with larger pit
on one hand or those without it on the other.
4) The dorsal pit with simple margin precedes that with double margins.
5) Nutlets with subbasal areola precede those with lateral or apical areola.
6) Nutlets without glochids precede those with glochids.
2. Basing upon these criteria the genus Microula may be divided into six sections.
The section Schistocaryum may be the primitive one, and the others may be evolved
from it respectively. The possible affinities between them are demonstrated in figure
no. two.
3. The genus Microula, containing 30 species, is mainly distributed in the
Chinghai-Tibetan plateau and the majority of its species concentrates in the eastern
border of the plateau, and of the 30 species 26—that is 90 percent—are endemic
to China, and the remaining 4 are distributed elsewhere in China, too, and extending
southward and westward to Bhutan, Sikkim, Nepal and Kashmir respectively. In the
region between Heishui, Province Szechuan, and Chinghai Lake there are 9 species,
which, curiously, represent all the six sections of Microula, hence this region seems to
be the center of maximum variation of this genus. M. ovalifolia whose nutlets have
small dorsal pit and subbasal areola may be considered the most primitive species.
Thus the author is of the opinion that the western part of province Szechuan, to
which M. ovalifolia is endemic, may probably be the center of origin of the genusMicroula. 相似文献
6.
1) The Compositae in Tibet so far known comprise 508 species and 88 genera,
which nearly amounts to one fourth of the total number of genera and one third of the
total number of species of Compositae in all China, if the number of 2290 species and 220
genera have respectively been counted in all China. In Tibet there are all tribes of Com-
positae known in China, and surprisingly, the large tribes in Tibetan Compositae are
also large ones in all China and the small tribes in Tibet are also small ones in all China.
Generally speaking, the large genera in Tibet are also large ones in all China and the
small genera in Tibet are likewise small ones in all China. In this sense it is reasonable to
say that the Compositae flora of Tibet is an epitome of the Compositae flora of all China.
In the Compositae flora of Tibet, there are only 5 large genera each containing 30
species or more. They are Aster, Artemisia, Senecio, Saussurea and Cremanthodium. And
5 genera each containing 10—29 species. They are Erigeron, Anaphalis, Leontopodium,
Ajania, Ligularia and Taraxacum. In addition, there are 77 small genera, namely 87%
of the total of Compositae genera in Tibet, each comprising 1—9 species, such as Aja-niopsis, Cavea and Vernonia, etc.
2) The constituents of Compositae flora in Tibet is very closely related to those of
Sichuan-Yunnan provinces with 59 genera and 250 species in common. Such a situation
is evidently brought about by the geographycal proximity in which the Hengtuang Shan
Range links southeastern and eastern Tibet with northern and northwestern Sichuan-
Ynnnan. With India the Tibetan Compositae have 59 genera and 132 species in common,
also showing close floristic relationships between the two regions. Apparently the floris-
tic exchange of Compositae between Tibet and India is realized by way of the mountain
range of the Himalayas. The mountain range of the Himalayas, including the parallel
ranges, plays a important role as a bridge hereby some members of the Compositae of
western or northern Central Asia and of the northern Africa or of western Asia have
migrated eastwards or southeastwards as far as the southern part of Fibet and northern
part of India, or hereby some Compositae plants of eastern and southeastern Asia or
Asia Media have migrated northwestwards as the northern part of Central Asia.
Some of the species and genera in common to both Tibet and Sinjiang indicate that
this weak floristical relationship between these regions is principally realized through two
migration routes: one migration route is by way of the Himalayas including the parallel
ranges to Pamir Plataeu and Tien Shan, or vice versa. The other migration route is by
way of northern Sinjiang to Mongolia, eastern Inner Mongolia, southwards to Gansu,
Qinghai (or western Sichuan), eastern Tibet up to the Himalayas, or vice versa.
However, Tibet is not entirely situated at a migration crossroad of the floral ele-
ments. An ample amount of the data shows that Compositae flora have a particular
capability of development in Tibet. of the total number of species of Tibetan Com-
positae, 102 species and 1 genus (Ajaniopsis Shih) are endemic. Besides, 8 genera are re-
gional endemics with their range extending to its neighbourhood. The higher percentage
of endemics at specific level than at generic in Tibetan Compositae may be a result of
active speciation in response to the new enviromental conditions created by the uplifting
of the Himalayas. The flora in Tibetan Plateau as a whole appears to be of a younger
age.
3) The uprising of the Himalayas and of the Tibetan Plateau accompanied by the
ultraviolet ray radiation, the microthermal climate and the high wind pressure has, no
doubt, played a profound influence upon the speciation of the native elements of Tibetan
Compositae. The recent speciation is the main trend in the development of the Com-positae flora native in Tibet in the wake of upheaval of the plateau. 相似文献
7.
We have described a new genus Taihangia, collected from, the south part of Taihang
Mountain in northern China. At the same time, comparative studies on Taihangia with its
related genera have been made in various fields including external morphology, anatomy
of carpels, chromosome and pollen morphology by light, scanning and transmission electron microscope. In addition, isoperoxidases of two varietier were analysed by means of polya-crylamide gel slab electrophoresis. The preliminary results are as follows:
Morphology: The genus Taihangia is perennial and has simple leaves, occasionally
with 1—2 very small reduced lobes on the upper part of petiole; flowers white, andromo-
noecious and androdioecious, terminal, single or rarely 2 on a leafless scape; calyx and cpicalyx with 5 segments; petals 5; stamens numerous; pistils numerous, with pubescent styles, spirally inserted on the receptacle in bisexual flowers, but with less number of abortive and glabrous pistils in male flowers.
In comparison with the related genera such as Dryas, Geum, Coluria and Waldsteinia,
the new genus has unisexual flowers and always herbaceous habit indicating its advanced
feature but the genus has a primitive style with thin and short hairs as compared with the genus Dryas which has long, pinnately haired styles, a character greatly facilitamg anemo-choric dissemination. The styles of Taihangia are slender and differ from those of the ge-nus Geum which are articulate, with a persistent hooked rostrum, thus adapting to epizo-ochoric dissemination to a higher degree.
The anatomy of carpels shows the baral position of ovules in the genus Taihangia like those in other related genera such as Dryas, Geum, Acomastylis, Coluria and Waldsteinia. This suggests that the new genus and its related ones are in a common evolutionary line as compared with the other tribes which have a pendulous ovule and represent a separate evolutionary line in Rosaceae. Dorsal and ventral bundles in carpels through sections are free at the base. Neither fusion, nor reduction of dorsals and vertrals. are observed. This shows that the genus Taihangia is rather primitive.
Somatic chromosome: All the living plants, collected from both Honan and Hopei
Provinces were examined. The results show that in these plants the chromosome number
is 2n= 14, and thus the basic number of chromosome is x=7. Such a diploid genus is first
found in both anemochoric and epizoochoric genera. Therefore, in this respect Taihangia is primitive as compared with herbaceous polyploid genus Geum and related ones.
Pollen: The stereostructure shown by scanning electron microscope reveals that the pollen grains of the genus Taihangia are ellipsoid and 3-colporate. There are two types of exine sculpture. One is rather shortly striate and it seems rugulate over the pollen surface; the other is long-striate. The genus Dryas differs in having only short and thick striae over the surface. The genus is similar to the genera Geum, Coluria and Waldsteinia in colpustype, but differs from them in that they all have long, parallel striae which are distributed along the meridional line.
In addition, under transmission electron microscope, the exine in the Taihangia and related genera Acomastylis, Geum, Coluria, Waldsteinia and Dryas has been shown to be typically differentiated into two distinct layers, nexine and sexine. The nexine, weakly statined, appears to consist of endoxine with no foot-layer, in which the columellae are fused, and which is thicker beneath the apertures. The sexine is 2-layered, consisting of columellae and tectum. Three patterns of tectum can be distinguished in the tribe Dryadeae: the first, in the genera Taihangia, Acomastylis, Geum, Coluria and Waldsteinia, is tectate-imperforate, with the sculpturing elements both acute and obtuse at the top and broad at the base; the second, in the genus Dryas, is semitectate, with the sculpturing elements shown in ultrathin sections rod-like and broader at the top than at the base or as broad at the top as at the base, and the third, tectate-perforate, with the sculpturing elements different in size. From the above results, the herbaceous groups and woody ones have palynologically evolved in two distinct directions, and the genus Taihangia is related to other herbaceous genera such as Acomastylis, Geum, Coluria and Waldsteinia, as shown in the electron microphotographs of ultrathin sections. The genus Taihangia, however, is different from related herbaceous genera in that the pollen of Taihangia is dimorphic, i.e. in addition to the above pattern of pollen another one of the exine in Taihangia is rugulate, with the sculpturing elements shown in the ultrathin sections being obtuse or emarginate and nearly as broad at the top as at the base.
The interesting results obtained from the comparative analysis of morphology, ana-
tomy of carpels, chromosome countings, microscopic and submicrosocopic structures of pollen may enable us to evaluate the systematic position of Taihangia and to throw a new light on evolution of the tribe Dryadeae. It is well known that the modes of dissemination of rosaceous fruits play an important role in the expansion and evolution of the family. The follicle is the most primitive and the plants with follicles, like the Spiraeoideae, are mostly woody and mesic, while the achene, drupe and pyrenarium are derived. In Rosoideae having a achene is a common feature. Particularly in the tribe Dryadeae, which is distinguished from the other related tribes by having orthotropous ovules, the methods of dissemination of fruits have developed in three distinct specialized directions: anemochory with long, plumose styles (e.g. Dryas), formicochory or dispersed by ants or other insects, with the deciduous styles (e.g. Waldsteinia and Collria),and epizoochory with the upper deciduous stigmatic part and the lower persistent hooked rostrum, an adhesive organ favouring epizoochory dissemination (e. g. Geum and related taxa). Taihangia is a genus endemic to mesophytic
forest area of northern China. Due to its narrow range and specific habit as well as pubescent styles, neither perfectly adapted to anemochory nor to epizoochory, the genus Taihangia might be a direct progeny of the ancestry of anemochory. Maintaining the diploidy and having an ntermediate sculptural type of pollen, the new genus might probably represent a linkage between anemochory and zoochory (including epizoochory and dispersed by ants).
Experimental evidence from isoperoxidases shows the stable zymograms of root and
roostoks. The anodal isozyme of T. rupestris var. rupestris may be divided into 6 bands:
A, B, C, D, E, F, and T. rupestris var. ciliata into 4 bands: A, B, C, G. The two varietiesof the species share 3 bands: A, B, C. However, D, E and F bands are characteristic of var. rupestris and G band is limited to var. ciliata. As far as the available materials are concerned, the analysis of isoperoxidases supports the subdivision of the species into two varieties. 相似文献
8.
9.
我国悬钩子属植物的研究 总被引：1，自引：0，他引：1
陆玲娣 《中国科学院研究生院学报》1983,21(1):13-25
The genus Rubus is one of the largest genera in the Rosaceae, consisting of more
than 750 species in many parts of the world, of which 194 species have been recorded
in China.
In the present paper the Rubus is understood in its broad sense, including all the
blackberries, dewberries and raspberries, comprising the woody and herbaceous kinds.
So it is botanically a polymorphic, variable and very complicated group of plants.
The detailed analysis and investigation of the evolutionary trends of the main organs
in this genus have indicated the passage from shrubs to herbs in an evolutionary line,
although there is no obvious discontinuity of morphological characters in various taxa.
From a phylogenetic point of view, the Sect. Idaeobatus Focke is the most primitive
group, characterized by its shrub habit armed with sharp prickles, aciculae or setae,
stipules attached to the petioles, flowers hermaphrodite and often in terminal or axill-
ary inflorescences, very rarely solitary, druplets separated from receptacles. Whereas
the herbaceous Sect. Chamaemorus L. is the most advanced group, which is usually
unarmed, rarely with aciculae or setae, stipules free, flowers dieocious, solitary, dru-
plets adhering to the receptacles and with high chromosome numbers (2n = 56).
Basing upon the evolutionary tendency of morphological features, chromosome nu-
mbers of certain species recorded in literature and the distribution patterns of species,
a new systematic arrangement of Chinese Rubus has been suggested by the present
authors. Focke in his well-known monograph divided the species of Rubus into 12
subgenera, while in the Flora of China 8 sections of Focke were adapted, but some im-
portant revisions have been made in some taxa and Sect. Dalibarda Focke has been
reduced to Sect. Cylactis Focke. In addition, the arrangement of sections is presented
in a reverse order to those of Focke’s system. The species of Rubus in China are
classified into 8 sections with 24 subsections (tab. 3) as follows: 1. Sect. Idaeobatus,
emend. Yü et Lu(11 subsect. 83 sp.); 2. Sect. Lampobatus Focke (1 sp.); 3. Sect.
Rubus (1 sp.); 4. Sect. Malachobatus Focke, emend. Yü et Lu (13 subsect. 85 sp.); 5.
Sect. Dalibardastrus (Focke)Yü et Lu (10 sp.); 6. Sect. Chaemaebatus Focke (5 sp.);
7. Sect. Cylactis Focke, emend. Yü et Lu (8 sp.); 8. Sect. Chamaemorus Focke (1 sp.).
In respect to the geographical distribution the genus Rubus occurs throughout the
world as shown in tab. 2, particularly abundant in the Northern Hemisphere, while
the greatest concentration of species appears in North America and E. Asia. Of the
more than 750 species in the world, 470 or more species (64%) distributed in North
America. It is clearly showm that the center of distribution lies in North America at
present time. There are about 200 species recorded in E. Asia, of which the species
in China (194) amount to 97% of the total number. By analysis of the distribution
of species in China the great majority of them inhabit the southern parts of the Yangtze
River where exist the greatest number of species and endemics, especially in south-
western parts of China, namely Yunnan, Sichuan and Guizhou (tab. 3. 4.). It is in-
teresting to note that the centre of distribution of Rubus in China ranges From north-
western Yunnan to south-western Sichuan (tab. 5), where the genus also reaches its
highest morphological diversity.
In this region the characteristics of floristic elements of Rubus can be summarized
as follows: it is very rich in composition, contaning 6 sections and 94 species, about
66% of the total number of Chinese species; there are also various complex groups,
including primitive, intermediate and advanced taxa of phylogenetic importance; the
proportion of endemic plants is rather high, reaching 61 species, up to 44% of the
total endemics in China. It is noteworthy to note that the most primitive Subsect.
Thyrsidaei (Focke) Yü et Lu, consisting of 9 endemic species, distributed in southern
slopes of the Mts. Qin Ling and Taihang Shan (Fig. 4). From the above facts we may
concluded that the south-western part of China is now not only the center of distribu-
tion and differentiation of Rubus in China, but it may also be the center of origin ofthis genus. 相似文献
10.
芍药属的研究(1)——国产几个野生种核型的报道 总被引：1，自引：0，他引：1
In the present paper 8 species with 15 populations of the genus Paeonia L. (if P.
papaveracea and P. japonica are recognised as species) were collected from Sichuan, Shaanxi
and Hebei provinces (see the Appendix for detail of the materials). The micrographs of their
somatic metaphase (also Mii in the case of P.veitchii) are shown in Plates 1-4, the karyo-
type formulae, ranges of chromosome length and classification of karyotypes according to Steb-
bins (1971) are shown in Table 5: the idiograms in Figs. 1-2, and the parameters of chromo-
somes in Table 1-4. The essential points are mentioned as follows:
(1) Chromosomes of the various species in the section Modan have so far been examined
and they are all diploid, the two species in the section Onaepia are also diploid, and thus
tetraploids exist only in the section Paeonia.
(2) Chromosomes in the genus Paeonia are relatively stable except for the differentiation
of ploidy. The karyotypes (Table 1-4) show no differences among different taxa in Sect.
Modan and the same can also be said about the taxa in Sect. Paeonia (Table 1). Not only
are the karyotypes very similar, but also among the members within either section have the
same parameters of chromosomes, and, differences, if occur, are not statistically significant.
Between the two sections, however, the situation is different. The arm ratios of the first pairs of
chromosomes in Sect. Modan are 1.53, 1.52 and 1.48 (Table 1), but those in Sect. Paeonia
are 1.12-1.28 (Table 2-4), 95% confidence limits are 1.46-1.60 for the section Modan and
1.07-1.28 (1.21-1.35 only for PB85078) for the section Paeonia, not overlapping, which indi-
cates that the two sections have differentiated in respect of the first pairs of chromosomes.
(3)The population PB85024, which belongs to the P. obovata complex, has a karyotype of
2B (stebbins, 1971), which is a new one in the genus Paeonia. This karyotype is a stable one,
for several individuals in the population are uniform in this respect, which shows that Steb-
bins’ (1971) generalization that all the species in Paeonia have 2A does not hold true.
(4) Three populations of P. obovata complex studied in this work from Sichuan and
Shaanxi are all tetraploids, and one from Hebei is a diploid. From the present work and
the previcus reports, the materials from Japan and Korea, no matter whether flowers are pink
or white, are diploids, those from Heilongjiang Province (with both pink and white flowers)
(Liu Ming-yuan, personal communication) and from Heibei Province (with pink flowers)
in China are also diploids, the one from Sakhalin (pink flowers) is tetraploid, those from
Priamur of the Soviet Union are a tetraploid (with white flowers) and a diploid (with pink
flowers), and those from Shaanxi (the Qinling Range) and western Sichuan (with both pink
and white flowers) are all tetraploids. As far as we have now known, ploidy in this parti-
cular complex is correlated with the geographical distribution: diploids are found in the cen-
tral part, tetraploids occur in the northern limits, and in the south letraploids are the only
cytotype.
(5) The materials of P. mairei from western Sichuan and Shaanxi (the Qinling Range)
are found all to be tetraploids, which shows that two cytotypes, diploid and tetraploid, exist in
this species, but the geographical distribution pattern of these two cytotypes is to be revealed
in the future investigation. 相似文献
11.
Cypripedium subtropicum S. C. Chen et K. Y. Lang is a phytogeography-
cally significant new species with its habit, inflorescence and column very similar to
those of Selenipedilum of tropical America. It is found in Mêdog of southeastern Xi-
zang, China. Its slender leafy stem bears at the summit a many-flowered raceme, am-
ounting to 1.5 m in height. Although its ovary is unilocular—this is the reason why
we place it in Cypripedium, the column characters resemble those of Selenipedilum. For
example, the staminode is rather small and its long stalk is very similar in texture and
color to the filament of the fertile stamens. Obviously, it is a primitive new species re-
lated to Selenipedilum based on the similarities mentioned above.
In the subfamily Cypripedioideae, as generally recognized, Selenipedilum is the
most primitive genus, from which or whose allies Cypripedium is derived. Of phyto-
geographical significance is the fact that Selenipedilum occurs in Central America and
northern South America, while a cypripedium akin to it is discontinuously distributed
in subtropical Asia. This suggests that Selenipedilum or Selenipedilum-like form be
once continually distributed in North America and eastern Asia when the climate there
was warmer, as it is in the subtropics today. The floristic relationship between Central
America and subtropical Asia appears to be closer than expected, as shown by the dis-
tribution patterns of Tropidia, Erythrodes, etc. Based on the occurrence of all six sec-
tions and particularly the most primitive form in eastern Asia, Cypripedium seems to
be of Asian, rather than Central American, origin. Selenipedilum possesses some very
primitive characters, such as trilocular ovary, vanilla-scented fruit, seed with sclerotic
testa, simple column and more or less suffrutescent habit. The latter is considered by
Dahlgren & Clifford (1982) to be one of ancestral characters of monocotyledons, which
is now very rare not only in Orchidaceae but also in all monocotyledons. It is indeed
necessary to make further investigations on Selenipedilum and also the new species pub-lished here, as well as a detailed comparison between them. 相似文献
12.
傅德志 《中国科学院研究生院学报》1988,26(4):249-264
本文对人字果属Dichocarpum W．T．Wang et Hsiao．的形态、花粉和染色体等性
状，以及地理分布进行了系统研究。确认了该属在毛茛科Ranunculaceae中的地位，并认为可
能与星果草属Asteropyrum Drumm．et Hutch．关系较密切, 证实了该属内存在三沟和散沟两
种花粉类型。该属的染色体基数可能为x=6，产于东亚大陆的种为4倍体，日本的种为6倍
体，原始的2倍体种已灭绝。中国西部山地可能为该属的分布中心，日本的种可能是在第三纪由中国大陆迁移过去的。本文按该属内各种之间可能的亲缘关系，作出了系统排列。 相似文献
13.
The classical and numerical taxonomy, palynology and the geographical dis-
tribution of the Genus Schizopepon are dealt with in the present paper. Having comme-
nted on various opinions regarding the systematic position of the genus, the present au-
thors consider that C. Jeffrey’s treatment of Schizopepon as a new and monogeneric tri-
be, Schizopeponeae, should be supported.
The gross morphological characters in the genus are assessed from the taxonomic
point of view. Some characters, such as stamens with an elongated connective or not,
different insertions of ovules and various forms of ovaries and fruits, may be used for
distinguishing subgenera.
The pollen grains of all the species were observed under light microscope (LM) and
scanning electron microscope (SEM). The results show that a strong differentiation
has taken place in the pollen of the genus, and in consequence it may be regarded as an
important basis for dividing subgenera and species. Especially it should be pointed out
that degrees of development of colpi and positions of ora are positively correlated with
the external characters used for distinguishing subgenera.
According to the morphological and palynological characters, the genus Schizopepon
may be divided into three subgenera and eight species: 1. Subgenus Schizopepon: 5 spe-
cies, S. bryoniaefolius Maxim., S. monoicus A. M. Lu et Z. Y. Zhang, S. dioicus Cogn.,
S. longipes Gagnep. and S. macranthus Hand.-Mazz.; 2. Subgenus Rhynchocarpos A. M.
Lu et Z. Y. Zhang: 1 species, S. bomiensis A. M. Lu et Z. Y. Zhang; 3. Subgenus Neoschi-
zopepon A. M. Lu et Z. Y. Zhang: 2 species, S. bicirrhosus (C. B. Clarke) C. Jeffrey
and S. xizangensis A. M. Lu et Z. Y. Zhang.
The 8 OTU’s including all the species of this genus and 31 characters, of which 16 are morphological characters and 15 palynological characters, were used in the numerical taxonomic treatment. After standardization of characters, the correlation and distance matrices were computed. The correlation matrices are made to test the various clustering methods. At last, the UPGMA clustering method was selected and its result
is shown in the form of phenogram. The result of numerical analysis is similar to that of the classical classification.
Schizopepon Maxim. is a genus of East Asia-Himalayan distribution. China has all 8 species and 2 varieties, of which 6 species are endemic. Based on the statistics of spedies number, the distribution centre of the genus is considered to be in the Hengduan Mountains (Yangtze-Mekong-Salwin water divides) and the adjacent areas of the southwest China. 相似文献
14.
15.
李恒 《中国科学院研究生院学报》1985,23(2):144-148
The present paper reports the first record of the eGnus Stachyphryni-
um, Marantaceae, in China. It is characterized by solitary spikes, elongated and erect
with impricated bracts. The inflorescence arises from a short stem on the rhizome.
In this genus there are 14 species in total. They are distributed from Sri Lanke
to Java and Borneo, through Indo-China Peninsula, Malay Pennisula. Its distribution
center is in Indo-China Penninsula and Malay Penninsula. The north limit lies in sou-
thern Yunnan of China. Most species of the genus have a stenochorie area, for example,
S. zeylanicum is endemic to Sri Lanka; S. latifolium occurs in Java only and so on.
A New species, S. Sinense H. Li, to China, is illustrated and described in Latin. 相似文献
16.
本文根据植物类群的系统发育和地理分布统一的原理，讨论了獐牙菜属植物的起源、散布和分
布区的形成。獐牙菜属包括11组16系154种，间断分布在亚洲、欧洲、北美洲和非洲。中国西南部-
喜马拉雅地区汇集了大多数种类、不同演化水平的类群以及形形色色的特有类群，成为该属的多样化
中心和多度中心。该属的原始类群和外类群也集中分布在中国西南山地，极有可能是该属的起源地。该
属的分布区类型中出现了各式的间断分布，根据有该属植物分布的大陆间及大陆与岛屿间分离和连接
的时间推测，该属的起源时间至少不会晚于晚白垩纪，也许更早，可追溯到中白垩纪。通过分类群间亲
缘关系和现代分布分析，显示出该属植物从起源地向周围和一定方向散布，形成了三个主要散布途径。在散布过程中植物本身也发生演化和就地特化，形成新的类群。 相似文献
17.
阴山荠属的校订 总被引：1，自引：0，他引：1
张渝华 《中国科学院研究生院学报》1987,25(3):204-219
The genus Yinshania was established by Ma Yu-chuan and Zhao Yi-zhi in 1979,
when only one species, Y. albiflora Ma et Y. Z. Zhao, was discribed from Nei Monggol. In
the present paper the genus Yinshania is revised and four new species, two new varieties and
four new combinetions are reported. There are so far eight species and two varieties in total
in this genus.
Important morphological characters of the genus are analysed, which shows that the lateral
nectariferous glands positioned at lateral base of the brevistamens are triangularovoid; there are
dense minute pustules on the surface of valves, which is easily neglected because the pustules disap-
pear or shrinked when dry; simple or furcate hairs are present in the most species, seldom absent;
the shape of pollen grains is relatively steady, elliptic or long-elliptic, with the polar view trifid-
circular, the equatorial view elliptic or long elliptic, the aperture 3-colpate, exine reticular.
The type of genus Yinshania is changed. Cochlearia acutangula O. E. Schulz was published
in 1929, but Y. albiflora Ma et Y. Z. Zhao in 1979. They are the same species and a new com-
binetion, Y. acutangula (O. E. Schulz) Y. H. Zhang, is made. Thus, the type of genus Yin-
shania should be changed to Y. acutangula (O. E. Schulz) Y. H. Zhang.
Besides, He Ye-qi 6121 (paratype, PE), which is different from Y. acutangula var. albif-
lora, is separated from it and transferred the typical variety, Y. acutangula.
According to the characters of fruit shape the genus Yinshania is divided into two sections,
namely, Sect. Microcarpa and Sect. Yinshania, and then Sect. Yinshania is subdivided into two
series.
Sect. 1. Microcarpa. Silicles widely ovoid or subglobose, 1-2.2 mm long, 0.8-2.2 mm wide,
the ratio of length and width about 1.1.
Sect. 2. Yinshania. Silicles oblong, oblong-ovoid or long-lanceolate, ellipsoidal, 1.5-4.5 mm
long, 0.3-1.5 mm wide, the ratio of length and width about 2.5-3.3.
Ser. 1. Henryanae. Raches flexuose; plants densely hairy; leaves 3-5-foliolate, seldom
pinnatipartite or pinnatisect.
Ser. 2. Yinshania. Raches non flexuose; plants sparsely hairy; leaves pinnatisect or pinna-
tipartite.
The genus Yinshania is a genus endemic to China, with their range from eastern Xizang to
western Hubei from northern Guizhou to central Nei Monggol. The taxa are mostly of a small
area. Sect. Microcarpa is concentrated in Sichuan and southern Gansu; Sect. Yinshania is
spread from Xizang and Sichuan, nouthwards to Gansu, Ningxia, Shanxi, Hebei and Nei Mong-
gol (Ser. Yinshania); and from Sichuan south-eastwards to Guizhou and Hebei (Ser. Henrya-
nae). There are five species in Sichuan. The present paper conjectures that the distribution
centre of the genus is in the Hengduan Mountains and its adjacent areas. 相似文献